Habituation and classical conditioning
Habituation and dishabituation
Habituation refers to the process by which an individual becomes so accustomed to a stimulus through prolonged or repeated exposure that it essentially fades into the background of awareness. Over time, one learns to disregard that stimulus. For example, when spending time in a crowded bar with loud music, the initial noise level gradually becomes unnoticeable as one adapts, so that later, when stepping outside into the relatively quiet sounds of a street or parking lot, the environment seems surprisingly muted.
Dishabituation is the reversal of this process, wherein the stimulus that was once ignored regains its prominence and captures attention once again. This reversal can occur after a brief or extended interval. In other words, if an individual who has become habituated to a particular environment suddenly encounters a change in that environment, the previously ignored stimulus can again become noticeable.
When considering behaviors, it is useful to distinguish between innate and learned actions. Innate behaviors include both instincts and reflexes—responses that are present at birth. Reflexes are motor or neural responses to specific environmental stimuli. They are typically simpler, involve the activation of particular body parts or systems (for instance, the knee-jerk reflex or the contraction of the pupil when exposed to bright light), and are controlled by more primitive areas of the central nervous system such as the spinal cord and medulla.
In contrast, instincts are broader patterns of behavior that are also unlearned but are triggered by a wider range of events, such as changes associated with maturation or seasonal variations. Instincts usually manifest as more complex behaviors involving the entire organism, including activities like sexual behavior or migration, and they rely on higher brain centers.
Learned behaviors, on the other hand, result from change and experience. Learning is defined as a relatively enduring change in behavior or knowledge that arises as a consequence of experience, differing from innate behaviors in that it involves the acquisition of new skills or information over time.
A key process underlying learning is associative learning, the natural tendency of the mind to connect events that occur in close temporal proximity or in a specific sequence. Through associative learning, an organism forms connections between stimuli or events that occur together in its environment. This process is fundamental to several types of learning, including classical conditioning.
In classical conditioning—also known as Pavlovian conditioning—organisms learn to associate events or stimuli that repeatedly occur together. For instance, during a storm, the sight of a flash of lightning is frequently followed by the sound of thunder. Over time, the sound of thunder, which naturally elicits a startle response, becomes associated with the flash of lightning, so that merely observing lightning may prompt a similar reaction.
In Pavlov’s experiments, the research involved measuring the salivary response of dogs to various stimuli. Initially, the presentation of meat powder naturally caused the dogs to salivate, an unlearned response termed the unconditioned response (UCR) elicited by an unconditioned stimulus (UCS). The dogs did not salivate in response to stimuli that were unrelated to food, such as an empty bowl or the sound of footsteps, because these stimuli were neutral (NS) prior to conditioning.
During conditioning, a neutral stimulus, such as a tone, was presented immediately before the unconditioned stimulus (meat powder).
Over repeated pairings, this neutral stimulus began to elicit salivation on its own, thereby transforming into a conditioned stimulus (CS) that produced a conditioned response (CR) identical to the original unconditioned response. In essence, the dogs learned to associate the tone with the arrival of food, leading them to salivate at the sound alone.

Furthermore, higher-order conditioning (or second-order conditioning) can occur when an additional stimulus is paired with the primary conditioned stimulus. This can happen intentionally in experiments or inadvertently, such as when a pet learns that the sound of a can opener predicts food, and then the sound of a squeaky cabinet, which consistently precedes the can opener’s noise, also becomes a secondary stimulus. However, achieving conditioning beyond second-order is generally challenging.
Discrimination and generalization
Another important aspect of learning is discrimination, which is the organism’s ability to distinguish between different stimuli. For example, an animal learns to respond only to a specific tone that reliably predicts food, while ignoring other similar sounds that do not have that association. In contrast, generalization occurs when an organism exhibits the conditioned response to stimuli that resemble the conditioned stimulus. The more similar a new stimulus is to the original conditioned stimulus, the more likely the conditioned response is to occur. However, through further learning, the organism may refine its responses to discriminate between stimuli if the differences become sufficiently distinct.
In summary, through these processes of habituation, dishabituation, innate and learned behaviors, and various forms of associative learning such as classical conditioning, organisms adapt to and interpret their environments, allowing them to respond appropriately to both familiar and novel stimuli.
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